For his PhD, Ryan is researching the phylogeography of 'giant' Collembola (primitive
insects - springtails - only these ones have no 'spring'!) and Symphyla (myriapods). These relatively unknown creatures
have important roles in decomposing wood (the
saproxylic habitat) although
we know relatively little about them.
In recognition of his excellent Honours year and numerous undergraduate awards, Ryan won the La Trobe University David
Myers PhD Scholarship. In 2003-5, he was awarded research grants by the ANZ Trustees, Holsworth Wildlife Research Fund,
and was sponsored to attend the International Congress of Genetics (Genetics Society of Australia) and 34th Australian
Entomological Society/ 6th Invertebrate Biodiversity & Conservation Conference (Department of the Environment and
Heritage – ABRS).
Ryan's work is based at our main comparative phylogeography study site at
Tallaganda
State Forest / National Park, NSW.
The results for the first species analysed are very striking. The 'smooth' springtail shows very strong differences
among regions within the forest, on the scale of a few Km. The network below left shows the evolutionary relationships
among different mitochondrial DNA sequences that can be found in smooth springtails in the forest. The frequencies
of these different types are represented as pie charts on the map to the right.
The different areas of the forest are thus thus home to different genetic types. While we are yet to see whether
these different types can be interpreted as separate species, they are certainly as genetically different as some other
organisms that do not naturally interbreed. Of particular interest is the area containing the yellow and green DNA
type and their sequences. This area is thought to represent a refuge (shelter) during the last glaciation, and indeed
harbours an important and diverse radiation of smooth springtails. A paper on this work is close to submission for
Molecular Ecology.

Penny with grand daughter Isabel
Collaborator
Penny Greenslade
Penny is working on the morphological taxonomy of the 'giant' Collembola we find at
Tallaganda The
cryptic species we find using genetic markers are especially interesting in the context of morphological taxonomy,
and through morphological assesment, it becomes feasible to understand the physical characters that are important in
the lives of the organisms we study and why these characters might vary from place to place